New to Cumberland in April 1961 as one of the first batch of FS's. It spent a lot of its time on country routes out of Whitehaven remaining on passenger carrying duties until June 1977 when along with 554 it became a driver trainer.

In 1983 , by which time it was the oldest vehicle in the fleet, it was decided to use 550 for publicity purposes and it was repainted into 1930's style livery. Despite this driver training was always its primary purpose especially after the demise of TV1 (ex 554) due to a cracked chassis frame. In 1990 its registration was transferred to another Cumberland vehicle and it was finally replaced as a trainer in 1991. Purchased in September of that year by the 550 Group it has since then been a regular attendee at rallies in the North of England having been repainted back into Tilling red and cream. In 2001 550 was once again reunited with its registration.